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singlet oxygen phototoxicity melanin oxidative stress photoreactivity skin PM2.5 eumelanin ARPE-19 EPR ROS aggregation free radicals keratinocytes pheomelanin rose bengal 18-DHN 2-methoxy-14- naphthoquinone BT-melanin BZ-melanin Bonds acceptors DHI-melanin DHICA-melanin Deep eutectic solvents Dopa-melanin Eosin Y HaCaT Hydrogen Hydrogen bonds donors MerTK receptor